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Rye (''Secale cereale'') is a
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
grown extensively as a
grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and legum ...
, a
cover crop In agriculture, cover crops are plants that are planted to cover the soil rather than for the purpose of being harvested. Cover crops manage soil erosion, soil fertility, soil quality, water, weeds, pests, diseases, biodiversity and wildlife i ...
and a
forage Forage is a plant material (mainly plant leaves and stems) eaten by grazing livestock. Historically, the term ''forage'' has meant only plants eaten by the animals directly as pasture, crop residue, or immature cereal crops, but it is also used m ...
crop. It is a member of the wheat
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...
(
Triticeae Triticeae is a botanical tribe within the subfamily Pooideae of grasses that includes genera with many domesticated species. Major crop genera found in this tribe include wheat (see wheat taxonomy), barley, and rye; crops in other genera include ...
) and is closely related to both
wheat Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeologi ...
(''Triticum'') and
barley Barley (''Hordeum vulgare''), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains, particularly in Eurasia as early as 10,000 years ago. Globally 70% of barley pr ...
(genus ''Hordeum''). Rye grain is used for
flour Flour is a powder made by grinding raw grains, roots, beans, nuts, or seeds. Flours are used to make many different foods. Cereal flour, particularly wheat flour, is the main ingredient of bread, which is a staple food for many culture ...
,
bread Bread is a staple food prepared from a dough of flour (usually wheat) and water, usually by baking. Throughout recorded history and around the world, it has been an important part of many cultures' diet. It is one of the oldest human-made f ...
,
beer Beer is one of the oldest and the most widely consumed type of alcoholic drink in the world, and the third most popular drink overall after water and tea. It is produced by the brewing and fermentation of starches, mainly derived from ce ...
,
crispbread Crispbread ( sv, knäckebröd (lit. crack bread), ''hårt bröd'' (hard bread), ''hårdbröd'', ''spisbröd'' (stove bread), ''knäcke'', da, knækbrød, no, knekkebrød, fi, näkkileipä or näkkäri, et, näkileib, is, hrökkbrauð, fo, k ...
, some whiskeys, some
vodka Vodka ( pl, wódka , russian: водка , sv, vodka ) is a clear distilled alcoholic beverage. Different varieties originated in Poland, Russia, and Sweden. Vodka is composed mainly of water and ethanol but sometimes with traces of impuritie ...
s, and animal
fodder Fodder (), also called provender (), is any agriculture, agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, domestic rabbit, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food g ...
. It can also be eaten whole, either as boiled rye berries or by being rolled, similar to
rolled oats Rolled oats are a type of lightly processed whole-grain food. Traditionally, they are made from oat groats that have been dehusked and steamed, before being ''rolled'' into flat flakes under heavy rollers and then stabilized by being lightly t ...
. Rye is a
cereal A cereal is any Poaceae, grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, Cereal germ, germ, and bran. Cereal Grain, grain crops are grown in greater quantit ...
grain and should not be confused with
ryegrass ''Lolium'' is a genus of tufted grasses in the bluegrass subfamily (Pooideae). It is often called ryegrass, but this term is sometimes used to refer to grasses in other genera. They are characterized by bunch-like growth habits. ''Lolium'' is ...
, which is used for lawns, pasture, and as hay for livestock.


Distribution and habitat

Rye is one of a number of species that grow wild in the
Levant The Levant () is an approximate historical geographical term referring to a large area in the Eastern Mediterranean region of Western Asia. In its narrowest sense, which is in use today in archaeology and other cultural contexts, it is eq ...
, central and eastern
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
and in adjacent areas. Evidence uncovered at the
Epipalaeolithic In archaeology, the Epipalaeolithic or Epipaleolithic (sometimes Epi-paleolithic etc.) is a period occurring between the Upper Paleolithic and Neolithic during the Stone Age. Mesolithic also falls between these two periods, and the two are someti ...
site of
Tell Abu Hureyra Tell Abu Hureyra ( ar, تل أبو هريرة) is a prehistoric archaeological site in the Upper Euphrates valley in Syria. The tell was inhabited between 13,000 and 9,000 years ago in two main phases: Abu Hureyra 1, dated to the Epipalaeolithi ...
in the
Euphrates The Euphrates () is the longest and one of the most historically important rivers of Western Asia. Tigris–Euphrates river system, Together with the Tigris, it is one of the two defining rivers of Mesopotamia ( ''the land between the rivers'') ...
valley of northern
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
suggests that rye was among the first
cereal A cereal is any Poaceae, grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, Cereal germ, germ, and bran. Cereal Grain, grain crops are grown in greater quantit ...
crops to be systematically cultivated, around 13,000 years ago. However, that claim remains controversial; critics point to inconsistencies in the
radiocarbon Carbon-14, C-14, or radiocarbon, is a radioactive isotope of carbon with an atomic nucleus containing 6 protons and 8 neutrons. Its presence in organic materials is the basis of the radiocarbon dating method pioneered by Willard Libby and coll ...
dates, and identifications based solely on grain, rather than on
chaff Chaff (; ) is the dry, scaly protective casing of the seeds of cereal grains or similar fine, dry, scaly plant material (such as scaly parts of flowers or finely chopped straw). Chaff is indigestible by humans, but livestock can eat it. In agri ...
. Domesticated rye occurs in small quantities at a number of
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several parts ...
sites in Asia Minor (
Anatolia Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...
, now Turkey), such as the
Pre-Pottery Neolithic B Pre-Pottery Neolithic B (PPNB) is part of the Pre-Pottery Neolithic, a Neolithic culture centered in upper Mesopotamia and the Levant, dating to years ago, that is, 8800–6500 BC. It was typed by British archaeologist Kathleen Kenyon during h ...
Can Hasan III near
Çatalhöyük Çatalhöyük (; also ''Çatal Höyük'' and ''Çatal Hüyük''; from Turkish ''çatal'' "fork" + ''höyük'' "tumulus") is a tell of a very large Neolithic and Chalcolithic proto-city settlement in southern Anatolia, which existed from appr ...
, but is otherwise absent from the archaeological record until the
Bronze Age The Bronze Age is a historic period, lasting approximately from 3300 BC to 1200 BC, characterized by the use of bronze, the presence of writing in some areas, and other early features of urban civilization. The Bronze Age is the second pri ...
of central Europe, c. 1800–1500 BCE. It is possible that rye traveled west from Asia Minor as a minor admixture in wheat (possibly as a result of
Vavilovian mimicry In plant biology, Vavilovian mimicry (also crop mimicry or weed mimicry) is a form of mimicry in plants where a weed evolves to share one or more characteristics with a domesticated plant through generations of artificial selection. It is name ...
), and was only later cultivated in its own right. Archeological evidence of this grain has been found in
Roman Roman or Romans most often refers to: *Rome, the capital city of Italy *Ancient Rome, Roman civilization from 8th century BC to 5th century AD *Roman people, the people of ancient Rome *''Epistle to the Romans'', shortened to ''Romans'', a letter ...
contexts along the
Rhine ), Surselva, Graubünden, Switzerland , source1_coordinates= , source1_elevation = , source2 = Rein Posteriur/Hinterrhein , source2_location = Paradies Glacier, Graubünden, Switzerland , source2_coordinates= , so ...
and the
Danube The Danube ( ; ) is a river that was once a long-standing frontier of the Roman Empire and today connects 10 European countries, running through their territories or being a border. Originating in Germany, the Danube flows southeast for , pa ...
and in Ireland and Britain. page 26. Roman naturalist
Pliny the Elder Gaius Plinius Secundus (AD 23/2479), called Pliny the Elder (), was a Roman author, naturalist and natural philosopher, and naval and army commander of the early Roman Empire, and a friend of the emperor Vespasian. He wrote the encyclopedic '' ...
was dismissive of a grain that may have been rye, writing that it "is a very poor food and only serves to avert starvation". He said it was mixed with
spelt Spelt (''Triticum spelta''), also known as dinkel wheat or hulled wheat, is a species of wheat that has been cultivated since approximately 5000 BC. Spelt was an important staple food in parts of Europe from the Bronze Age to medieval times. No ...
"to mitigate its bitter taste, and even then is most unpleasant to the stomach". Winter rye is any breed of rye planted in the autumn to provide ground cover for the winter. It grows during warmer days of the winter when sunlight temporarily warms the plant above freezing, even while there is general snow cover. It can be used to prevent the growth of
winter-hardy Hardiness of plants describes their ability to survive adverse growing conditions. It is usually limited to discussions of climatic adversity. Thus a plant's ability to tolerate cold, heat, drought, flooding, or wind are typically considered meas ...
weeds, and can either be harvested as a bonus crop or tilled directly into the ground in spring to provide more organic matter for the next summer's crop. It is sometimes used in
winter garden A winter garden is a kind of garden maintained in wintertime. History The origin of the winter garden dates back to the 17th to 19th centuries where European nobility would construct large conservatories that would house tropical and subtro ...
s and is a common
nurse crop Nurse crops are a subtype of nurse plants, facilitating the growth of other species of plants. The term is used primarily in agriculture, but also in forestry. Cover crops are a type of nurse crop. Agriculture In agriculture, a nurse crop is ...
. Rye grows better than any other cereal in heavy clay and light sandy and infertile or drought-affected soils. It can tolerate pH between 4.5 and 8.0, but soils having pH 5.0 to 7.0 are best suited for rye cultivation. Rye grows best in fertile, well-drained loam or clay-loam soils.


Frost resistance

''S. cereale'' can thrive in subzero environments. The leaves of winter rye produce various antifreeze polypeptides (different from the antifreeze polypeptides produced by some fish and insects).


Ecology

The nematode ''
Ditylenchus dipsaci ''Ditylenchus dipsaci'' is a plant pathogenic nematode that primarily infects onion and garlic. It is commonly known as the stem nematode, the stem and bulb eelworm, or onion bloat (in the United Kingdom).
'',
leaf beetle The insects of the beetle family Chrysomelidae are commonly known as leaf beetles, and include over 37,000 (and probably at least 50,000) species in more than 2,500 genera, making up one of the largest and most commonly encountered of all beetle ...
, fruit fly,
gout fly ''Chlorops pumilionis'' is a species of pest fly from the family Chloropidae. It is also known as the chloropid gout fly or barley gout fly. It is an oligophagous pest of cereal crops. Description The thorax is straw yellow, with three longitu ...
, cereal chafer, dart moth, cereal bug,
Hessian fly The Hessian fly or barley midge, ''Mayetiola destructor'', is a species of fly that is a significant pest of cereal crops including wheat, barley and rye. Though a native of Asia, upon its discovery it was believed to have been transported into ...
, and
rustic shoulder knot ''Apamea sordens'', the rustic shoulder-knot or bordered apamea, is a moth of the family Noctuidae. The species was first described by Johann Siegfried Hufnagel in 1766. It is distributed throughout Europe, east across the Palearctic to Central ...
are among insects which can seriously affect rye health.


Diseases

Rye is highly susceptible to the
ergot Ergot ( ) or ergot fungi refers to a group of fungi of the genus ''Claviceps''. The most prominent member of this group is ''Claviceps purpurea'' ("rye ergot fungus"). This fungus grows on rye and related plants, and produces alkaloids that ca ...
fungus. Consumption of ergot-infected rye by humans and animals results in a serious medical condition known as
ergotism Ergotism (pron. ) is the effect of long-term ergot poisoning, traditionally due to the ingestion of the alkaloids produced by the ''Claviceps purpurea'' fungus—from the Latin "club" or clavus "nail" and for "head", i.e. the purple club-head ...
. Ergotism can cause both physical and mental harm, including convulsions, miscarriage, necrosis of digits, hallucinations and death. Historically, damp northern countries that have depended on rye as a staple crop were subject to periodic epidemics of this condition. Such epidemics have been found to correlate with periods of frequent
witch trials A witch-hunt, or a witch purge, is a search for people who have been labeled witches or a search for evidence of witchcraft. The classical period of witch-hunts in Early Modern Europe and Colonial America took place in the Early Modern per ...
, such as the
Salem witch trials The Salem witch trials were a series of hearings and prosecutions of people accused of witchcraft in colonial Massachusetts between February 1692 and May 1693. More than 200 people were accused. Thirty people were found guilty, 19 of whom w ...
in Massachusetts in 1692. Modern grain-cleaning and milling methods have practically eliminated the disease, but contaminated flour may end up in bread and other food products if the ergot is not removed before milling. After an absence of 60 years,
stem rust Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, durum w ...
(''Puccinia graminis'' f. sp. ''tritici'') has returned to
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
.Multiple sources: Areas affected include
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
,
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
(
Western Siberia Western Siberia or West Siberia (russian: Западная Сибирь, Zapadnaya Sibir'; kk, Батыс Сібір) is a part of the larger region of Siberia that is mostly located in the Russian Federation. It lies between the Ural region an ...
),
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
, and
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
. Races
PTGBH Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, durum wh ...
,
PTGFH Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, durum w ...
,
RKBBK Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, durum wh ...
,
SKGBC Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, Durum, du ...
, SKGBH,
SKGBR Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, durum w ...
,
TKDLK Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, durum wh ...
,
TKGBK Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, durum wh ...
,
TKGBR Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, Durum, du ...
, and
TKGDH Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, durum wh ...
are now present on rye in Spain.


Invasiveness

Rye is a common, unwanted invader of winter wheat fields. If allowed to grow and mature, it may cause substantially reduced prices (docking) for harvested wheat.


Cultivation

Since the
Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the post-classical period of global history. It began with the fall of the Western Roman Empire a ...
, people have cultivated rye widely in
Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known as ...
and
Eastern Europe Eastern Europe is a subregion of the Europe, European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural, and socio-economic connotations. The vast majority of the region is covered by Russ ...
. It serves as the main
bread Bread is a staple food prepared from a dough of flour (usually wheat) and water, usually by baking. Throughout recorded history and around the world, it has been an important part of many cultures' diet. It is one of the oldest human-made f ...
cereal in most areas east of the
France–Germany border The Franco-German border (; ) separates France and Germany and has a length of , about half of it along the Rhine. History The Franco-German border can be traced back to the 17th century, and the various treaties following the Thirty Years' War ( ...
and north of
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
. In Southern Europe, it was cultivated on marginal lands. Rye grows well in much poorer soils than those necessary for most cereal grains. Thus, it is an especially valuable crop in regions where the soil has
sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural class of s ...
or
peat Peat (), also known as turf (), is an accumulation of partially decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, moors, or muskegs. The peatland ecosystem covers and is the most efficien ...
. Rye plants withstand cold better than other small grains do. Rye will survive with snow cover that would otherwise result in winter-kill for winter wheat. Most
farmer A farmer is a person engaged in agriculture, raising living organisms for food or raw materials. The term usually applies to people who do some combination of raising field crops, orchards, vineyards, poultry, or other livestock. A farmer mig ...
s grow winter ryes, which are planted and begin to grow in autumn. In spring, the plants develop and produce their crop. Autumn-planted rye shows fast growth; by the
summer solstice The summer solstice, also called the estival solstice or midsummer, occurs when one of Earth's poles has its maximum tilt toward the Sun. It happens twice yearly, once in each hemisphere ( Northern and Southern). For that hemisphere, the summer ...
, plants reach their maximum height of about a , while spring-planted wheat has only recently germinated. Vigorous growth suppresses even the most noxious weed competitors and rye can be grown without application of herbicides. Physical properties of rye affect attributes of the final food product such as seed size and surface area, and porosity. The surface area of the seed directly correlates to the drying and heat transfer time.Jouki, Mohammad; Emam-Djomeh, Zahra; and Khazaei, Naimeh (2012) "Physical Properties of Whole Rye Seed (Secale cereal)," International Journal of Food Engineering: Vol. 8: Iss. 4, Article 7. Smaller seeds have increased heat transfer, which leads to lower drying time. Seeds with lower amounts of porosity also have lower tendencies to lose water during the process of drying.


Diversity and uses

Along with ''S. cereale'''s relationship and impact on the environment, it is also a valuable species because of its expansive diversity and uses. In northern Portugal, fourteen different populations of ''S. cereale'' were analyzed in order to better understand their differences. It was discovered that the storage proteins are very diverse and possess a lot of overall genetic variation as well, which is useful information to know because scientists can use its diversity in breeding to produce the most efficient
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
of ''S. cereale''. Moreover, the beneficial characteristics of ''S. cereale'' can also be used to improve certain characteristics of other useful plants, like wheat. The pollination abilities of wheat were vastly improved when there was cross-pollination with ''S. cereale''. The addition of the rye chromosome 4R increased the size of the wheat anther along with increasing the number of pollen grains present. Along with improved wheat, the optimal characteristics of ''S. cereale'' can also be combined with another perennial rye, specifically ''S. montanum Guss'', in order to produce ''S. cereanum'', which has the beneficial characteristics of each. The hybrid rye (''S. cereanum'') can be grown in all environments, even with less than favorable soil and protects some soils from erosion. In addition, the plant mixture has improved forage and is known to contain digestible fiber and protein. Information about the diversity, the genome and ''S. cereanum'''s ability to cross fertilize with other species is useful information for scientists to know as they attempt to come up with various plant species that will be able to feed humanity in the future without leaving a negative footprint on the environment.


Harvesting

The
harvest Harvesting is the process of gathering a ripe crop from the fields. Reaping is the cutting of grain or pulse for harvest, typically using a scythe, sickle, or reaper. On smaller farms with minimal mechanization, harvesting is the most labor-i ...
ing of rye is similar to that of wheat. It is usually done with
combine harvester The modern combine harvester, or simply combine, is a versatile machine designed to efficiently harvest a variety of grain crops. The name derives from its combining four separate harvesting operations—reaping, threshing, gathering, and winnow ...
s, which cut the plants,
thresh Thresh may refer to: *Threshing, in agriculture **Threshing machine * A minor character in the novel ''The Hunger Games'' and its film adaptation * Thresh (gamer), handle of esports player Dennis Fong See also * Thrash (disambiguation) Thrash m ...
and winnow the grain, and release the
straw Straw is an agricultural byproduct consisting of the dry stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the yield of cereal crops such as barley, oats, rice, rye and wheat. It has a number ...
to the field where it is later pressed into bales or left as soil amendment. The resultant grain is stored in local
silo A silo (from the Greek σιρός – ''siros'', "pit for holding grain") is a structure for storing bulk materials. Silos are used in agriculture to store fermented feed known as silage, not to be confused with a grain bin, which is used t ...
s or transported to regional
grain elevator A grain elevator is a facility designed to stockpile or store grain. In the grain trade, the term "grain elevator" also describes a tower containing a bucket elevator or a pneumatic conveyor, which scoops up grain from a lower level and deposits ...
s and combined with other lots for storage and distant shipment. Before the era of
mechanised agriculture Mechanised agriculture or agricultural mechanization is the use of machinery and equipment, ranging from simple and basic hand tools to more sophisticated, motorized equipment and machinery, to perform agricultural operations. In modern times, po ...
, rye harvesting was a
manual Manual may refer to: Instructions * User guide * Owner's manual * Instruction manual (gaming) * Online help Other uses * Manual (music), a keyboard, as for an organ * Manual (band) * Manual transmission * Manual, a bicycle technique similar to ...
task performed with
scythe A scythe ( ) is an agricultural hand tool for mowing grass or harvesting crops. It is historically used to cut down or reap edible grains, before the process of threshing. The scythe has been largely replaced by horse-drawn and then tractor m ...
s or
sickle A sickle, bagging hook, reaping-hook or grasshook is a single-handed agricultural tool designed with variously curved blades and typically used for harvesting, or reaping, grain crops or cutting succulent forage chiefly for feeding livestock, ei ...
s. The cut rye was often shocked for drying or storage, and the threshing was done by manually beating the seed heads against a floor or other object.


Production and consumption statistics

Rye is grown primarily in Eastern, Central and Northern
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
. The main rye belt stretches from northern
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
through
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...
,
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
,
Belarus Belarus,, , ; alternatively and formerly known as Byelorussia (from Russian ). officially the Republic of Belarus,; rus, Республика Беларусь, Respublika Belarus. is a landlocked country in Eastern Europe. It is bordered by R ...
,
Lithuania Lithuania (; lt, Lietuva ), officially the Republic of Lithuania ( lt, Lietuvos Respublika, links=no ), is a country in the Baltic region of Europe. It is one of three Baltic states and lies on the eastern shore of the Baltic Sea. Lithuania ...
and
Latvia Latvia ( or ; lv, Latvija ; ltg, Latveja; liv, Leţmō), officially the Republic of Latvia ( lv, Latvijas Republika, links=no, ltg, Latvejas Republika, links=no, liv, Leţmō Vabāmō, links=no), is a country in the Baltic region of ...
into central and northern
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
. Rye is also grown in North America (
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
and the United States), in South America (
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
and
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
), in Oceania (
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
and
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
), in Turkey, in
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
and in northern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. Production levels of rye have fallen since 1992 in most of the producing nations, . For instance, production of rye in Russia fell from in 1992 to in 2020. Corresponding figures for other countries are as follows: Polandfalling from in 1992 to in 2020; Belarusfalling from in 1992 to in 2020; Chinafalling from in 1992 to in 2020. However, production levels rose in Germany from in 1992 to in 2020. World trade of rye is low compared with other grains such as wheat. The total export of rye for 2016 was $186M compared with $30.1B for wheat. Poland consumes the most rye per person at per capita (2009). Nordic and Baltic countries are also very high. The EU in general is around per capita. The entire world only consumes per capita.


Uses

Rye grain is refined into a
flour Flour is a powder made by grinding raw grains, roots, beans, nuts, or seeds. Flours are used to make many different foods. Cereal flour, particularly wheat flour, is the main ingredient of bread, which is a staple food for many culture ...
high in
gliadin Gliadin (a type of prolamin) is a class of proteins present in wheat and several other cereals within the grass genus ''Triticum''. Gliadins, which are a component of gluten, are essential for giving bread the ability to rise properly during baki ...
but low in
glutenin Glutenin (a type of glutelin) is a major protein within wheat flour, making up 47% of the total protein content. The glutenins are protein aggregates of high-molecular-mass (HMW) and low-molecular-mass (LMW) subunits with molar masses from about ...
. It therefore has a lower
gluten Gluten is a structural protein naturally found in certain cereal grains. Although "gluten" often only refers to wheat proteins, in medical literature it refers to the combination of prolamin and glutelin proteins naturally occurring in all grain ...
content than
wheat Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeologi ...
flour. It also contains a higher proportion of
soluble fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...
.
Alkylresorcinol Alkylresorcinols, also known as resorcinolic lipids, are phenolic lipids composed of long aliphatic chains and resorcinol-type phenolic rings. Natural sources of alkylresorcinols Alkylresorcinols are relatively rare in nature, with the main know ...
s are phenolic lipids present in high amounts in the bran layer (e.g.
pericarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggre ...
, testa and
aleurone Aleurone (from Greek ''aleuron'', flour) is a protein found in protein granules of maturing seeds and tubers. The term also describes one of the two major cell types of the endosperm, the aleurone layer. The aleurone layer is the outermost layer of ...
layers) of wheat and rye (0.1–0.3% of dry weight).
Rye bread Rye bread is a type of bread made with various proportions of flour from rye grain. It can be light or dark in color, depending on the type of flour used and the addition of coloring agents, and is typically denser than bread made from wheat f ...
, including
pumpernickel Pumpernickel (; ) is a typically heavy, slightly sweet rye bread traditionally made with sourdough starter and coarsely ground rye. It is sometimes made with a combination of rye flour and whole rye grains ("rye berries"). At one time it was ...
, is made using rye flour and is a widely eaten food in Northern and Eastern Europe. Rye is also used to make
crisp bread Crispbread ( sv, knäckebröd (lit. crack bread), ''hårt bröd'' (hard bread), ''hårdbröd'', ''spisbröd'' (stove bread), ''knäcke'', da, knækbrød, no, knekkebrød, fi, näkkileipä or näkkäri, et, näkileib, is, hrökkbrauð, fo, k ...
. Rye grain is used to make alcoholic drinks, such as
rye whiskey Rye whiskey can refer to two different, but related, types of whiskey: * American rye whiskey, which is similar to bourbon whiskey, but must be distilled from at least 51 percent rye grain * Canadian whisky, which is often referred to as (and o ...
and
rye beer Rye beer is a beer in which rye is substituted for some portion of the malted barley. ''Roggenbier'' is a beer produced with up to 60% rye malt. The style originated in Bavaria, southern Germany, and is brewed with the same type of yeast as a ...
. Other uses of rye grain include
kvass Kvass is a fermented cereal-based Alcohol by volume, low alcoholic beverage with a slightly cloudy appearance, light-brown colour and sweet-sour taste. It may be flavoured with berries, fruits, herbs or honey. Kvass stems from the northeastern ...
and an
herbal medicine Herbal medicine (also herbalism) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. With worldwide research into pharmacology, some herbal medicines have been translated into modern remed ...
known as rye extract. Rye
straw Straw is an agricultural byproduct consisting of the dry stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the yield of cereal crops such as barley, oats, rice, rye and wheat. It has a number ...
is used as
livestock bedding Bedding, in ethology and animal husbandry, is material, usually organic, used by animals to support their bodies when resting or otherwise stationary. It reduces pressure on skin, heat loss, and contamination by waste produced by an animal or thos ...
, as a
cover crop In agriculture, cover crops are plants that are planted to cover the soil rather than for the purpose of being harvested. Cover crops manage soil erosion, soil fertility, soil quality, water, weeds, pests, diseases, biodiversity and wildlife i ...
and
green manure In agriculture, a green manure is a crop specifically produced to be incorporated into the soil while still green. Typically, the green manure's biomass is incorporated with a plow or disk, as is often done with (brown) manure. The primary goal ...
for
soil amendment A soil conditioner is a product which is added to soil to improve the soil’s physical qualities, usually its fertility (ability to provide nutrition for plants) and sometimes its mechanics. In general usage, the term "soil conditioner" is often ...
, and to make
craft A craft or trade is a pastime or an occupation that requires particular skills and knowledge of skilled work. In a historical sense, particularly the Middle Ages and earlier, the term is usually applied to people occupied in small scale prod ...
s such as
corn dollies Corn dollies or corn mothers are a form of straw work made as part of harvest customs of Europe before mechanization. Before Christianisation, in traditional pagan European culture it was believed that the spirit of the corn (in American English, ...
. Rye flour is used in the original way to make
Falun red Falu red or falun red ( ; sv, falu rödfärg, ) is a permeable red paint commonly used on wooden cottages and barns in Sweden, Finland, and Norway. History Following hundreds of years of mining in Falun, large piles of tailings, residual prod ...
paint (in addition to linseed oil and iron oxide) in Sweden. Rye grain (aka "Rye Berries") is a popular medium to use as a grain spawn when cultivating some varieties of edible mushrooms. The grain is cleaned, hydrated, and sterilized and then injected with mushroom spores and the
mycelium Mycelium (plural mycelia) is a root-like structure of a fungus consisting of a mass of branching, thread-like hyphae. Fungal colonies composed of mycelium are found in and on soil and many other substrate (biology), substrates. A typical single ...
grow using the grain to obtain water and nutrients.


Food


Nutrition

A reference serving of rye provides of
food energy Food energy is chemical energy that animals (including humans) derive from their food to sustain their metabolism, including their muscle, muscular activity. Most animals derive most of their energy from aerobic respiration, namely combining the ...
and is a rich source (20% or more of the
Daily Value The Reference Daily Intake (RDI) used in nutrition labeling on food and dietary supplement products in the U.S. and Canada is the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97–98% of health ...
V of
essential nutrients A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...
, including
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
,
dietary fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...
, the
B vitamins B vitamins are a class of water-soluble vitamins that play important roles in cell metabolism and synthesis of red blood cells. Though these vitamins share similar names (B1, B2, B3, etc.), they are chemically distinct compounds that often coexist ...
,
niacin Niacin, also known as nicotinic acid, is an organic compound and a form of vitamin B3, an essential human nutrient. It can be manufactured by plants and animals from the amino acid tryptophan. Niacin is obtained in the diet from a variet ...
(27% DV) and vitamin B6 (23% DV), and several
dietary minerals In the context of nutrition, a mineral is a chemical element required as an essential nutrient by organisms to perform functions necessary for life. However, the four major structural elements in the human body by weight (oxygen, hydrogen, carbon, ...
(table). Highest nutrient contents are for
manganese Manganese is a chemical element with the symbol Mn and atomic number 25. It is a hard, brittle, silvery metal, often found in minerals in combination with iron. Manganese is a transition metal with a multifaceted array of industrial alloy use ...
(143% DV) and
phosphorus Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ear ...
(47% DV).


Health effects

According to
Health Canada Health Canada (HC; french: Santé Canada, SC)Health Canada is the applied title under the Federal Identity Program; the legal title is Department of Health (). is the Structure of the Canadian federal government#Departments, with subsidiary unit ...
and the U.S.
Food and Drug Administration The United States Food and Drug Administration (FDA or US FDA) is a List of United States federal agencies, federal agency of the United States Department of Health and Human Services, Department of Health and Human Services. The FDA is respon ...
, consuming at least per day of rye
beta-glucan Beta-glucans, β-glucans comprise a group of β-D-glucose polysaccharides ( glucans) naturally occurring in the cell walls of cereals, bacteria, and fungi, with significantly differing physicochemical properties dependent on source. Typically, ...
or per serving of
soluble fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...
can lower levels of
blood cholesterol Blood lipids (or blood fats) are lipids in the blood, either free or bound to other molecules. They are mostly transported in a protein capsule, and the density of the lipids and type of protein determines the fate of the particle and its influence ...
, a risk factor for
cardiovascular diseases Cardiovascular disease (CVD) is a class of diseases that involve the heart or blood vessels. CVD includes coronary artery diseases (CAD) such as angina and myocardial infarction (commonly known as a heart attack). Other CVDs include stroke, ...
. Eating whole-grain rye, as well as other high-fibre grains, improves regulation of
blood sugar Glycaemia, also known as blood sugar level, blood sugar concentration, or blood glucose level is the measure of glucose concentrated in the blood of humans or other animals. Approximately 4 grams of glucose, a simple sugar, is present in the blo ...
(i.e., reduces blood glucose response to a meal). Consuming
breakfast cereal Cereal, formally termed breakfast cereal (and further categorized as cold cereal or warm cereal), is a traditional breakfast food made from processed cereal grains. It is traditionally eaten as part of breakfast, or a snack food, primarily in ...
s containing rye over weeks to months also improved cholesterol levels and glucose regulation.


Health concerns

Like wheat,
barley Barley (''Hordeum vulgare''), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains, particularly in Eurasia as early as 10,000 years ago. Globally 70% of barley pr ...
, and their hybrids and derivatives, rye contains gluten, which makes it an unsuitable grain for consumption by people with
gluten-related disorders Gluten-related disorders is the term for the diseases triggered by gluten, including celiac disease (CD), non-celiac gluten sensitivity (NCGS), gluten ataxia, dermatitis herpetiformis (DH) and wheat allergy. The umbrella category has also been r ...
, such as
celiac disease Coeliac disease (British English) or celiac disease (American English) is a long-term autoimmune disorder, primarily affecting the small intestine, where individuals develop intolerance to gluten, present in foods such as wheat, rye and barl ...
,
non-celiac gluten sensitivity Non-celiac gluten sensitivity (NCGS) or gluten sensitivity is "a clinical entity induced by the ingestion of gluten leading to intestinal and/or extraintestinal symptoms that improve once the gluten-containing foodstuff is removed from the diet, a ...
, and
wheat allergy Wheat allergy is an allergy to wheat which typically presents itself as a food allergy, but can also be a contact allergy resulting from occupational exposure. Like all allergies, wheat allergy involves immunoglobulin E and mast cell response. ...
, among others. Nevertheless, some wheat allergy patients can tolerate rye or barley.
Ergotism Ergotism (pron. ) is the effect of long-term ergot poisoning, traditionally due to the ingestion of the alkaloids produced by the ''Claviceps purpurea'' fungus—from the Latin "club" or clavus "nail" and for "head", i.e. the purple club-head ...
is an illness that can result from eating rye and other grains infected by ergot fungi (which produce
ergoline Ergoline is a chemical compound whose structural skeleton is contained in a variety of alkaloids, referred to as ergoline derivatives or ergoline alkaloids. Ergoline alkaloids, one being ergine, were initially characterized in ergot. Some of thes ...
toxins in infected products). Although it is no longer a common illness because of modern
food safety Food safety (or food hygiene) is used as a scientific method/discipline describing handling, preparation, and storage of food in ways that prevent food-borne illness. The occurrence of two or more cases of a similar illness resulting from t ...
efforts, it was common before the 20th century, and it can still happen today if food safety vigilance breaks down.


Hybridization

Rye has long been considered an inferior grain to wheat in quality and digestibility, but has far larger kernels and is more hardy. In the 19th century, efforts were made to create a hybrid with the best qualities of both, known initially as triticosecale, but eventually becoming known as
triticale Triticale (; × ''Triticosecale'') is a hybrid of wheat (''Triticum'') and rye (''Secale'') first bred in laboratories during the late 19th century in Scotland and Germany. Commercially available triticale is almost always a second-generation h ...
. Initially fraught with fertility and germination problems, triticale is becoming more common worldwide in the 21st century, with millions of acres being produced.


Gallery


References


Further reading

* Schlegel provides
2011 updated version
online.


External links

* * * {{Authority control Cereals Plants described in 1753 Pooideae Staple foods